School of Biological Sciences, Life Sciences Building, University of Liverpool, Liverpool L69 7ZB, UK1
Biology Department, Dalhousie University, Halifax, Nova Scotia B3H 4J1, Canada2
For correspondence. Alan McCarthy. Tel: +44 151 794 4413 Fax: +44 151 794 4401. e-mail: aj55m{at}liverpool.ac.uk
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